On Macrocolura and the subdivision of Colura (Lejeuneaceae, Hepaticae)

Some newly recognized differences support the generic segregation of Macrocolura R.M.Schust. from Colura (Dumort.) Dumort. and are summarized in a key. Only a single species of Macrocolura is recognized: M. sagittistipula (Spruce) R.M.Schust. [= M. clavigera (Gottsche ex Ast) R.M.Schust., syn. nov.; = M. lyrata (Steph.) R.M.Schust., syn. nov.], which is described and illustrated. Based on differences presented in a key Colura is newly subdivided into two subgenera (Glotta Grolle & R.L.Zhu and Colura), of which the former comprises the sections Glotta Grolle & R.L.Zhu and Heterophyllum Ast and the latter the sections Colura, Gamolepis Ast, Harmophyllum Grolle and Oidocorys Ast ex Grolle. For ease of access the 25 species published since the Colura monograph by Jovet-Ast (1954a+b) are listed in alphabetical order with full references under the appropriate sections.